Tropical landscape is everything but stable. We are studying a number of drastic landscape changes that have occurred in the tropics over the past 140 million years and how they have influcence the extinction and origination of tropical biotas: from the extreme effects of global warming during the early Cenozoic to the global cooling of the Pliocene; from the lifting the Andes mountains to the creation of savannas; from periods with low levels of CO2 to events with extreme high levels of CO2. We use the geological, fossil and DNA records to answer these questions.